About the Authors

Zhihui Sun

Contributed equally to this work with: Zhihui Sun, Zamia Diaz

Affiliation Department of Cell and Developmental Biology, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Zamia Diaz

Contributed equally to this work with: Zhihui Sun, Zamia Diaz

Affiliation Department of Biochemistry and Biophysics, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Xiaodong Fang

Affiliation Department of Cell and Developmental Biology, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Michael P. Hart

Affiliation Department of Cell and Developmental Biology, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Alessandra Chesi

Affiliation Department of Cell and Developmental Biology, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

James Shorter

gitler@mail.med.upenn.edu (ADG); jshorter@mail.med.upenn.edu (JS)

Affiliation Department of Biochemistry and Biophysics, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Aaron D. Gitler

gitler@mail.med.upenn.edu (ADG); jshorter@mail.med.upenn.edu (JS)

Affiliation Department of Cell and Developmental Biology, The University of Pennsylvania School of Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: ZS ZD XF MPH JS ADG. Performed the experiments: ZS ZD XF MPH JS ADG. Analyzed the data: ZS ZD XF MPH AC JS ADG. Wrote the paper: JS ADG.