About the Authors

Kazuya Ono

Affiliations Sensory Development, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan, Neurogenesis and Organogenesis, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Tomoko Kita

Affiliation Sensory Development, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Shigeru Sato

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Paul O'Neill

Affiliation Sensory Development, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Siu-Shan Mak

Affiliation Sensory Development, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Marie Paschaki

Affiliation Sensory Development, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Masataka Ito

Affiliation Department of Anatomy, National Defense Medical College, Tokorozawa, Japan

Noriko Gotoh

Affiliation Division of Genetics, Institute of Medical Science, University of Tokyo, Minato-ku, Japan

Kiyoshi Kawakami

Affiliation Division of Biology, Center for Molecular Medicine, Jichi Medical University, Shimotsuke, Tochigi, Japan

Yoshiki Sasai

Affiliation Neurogenesis and Organogenesis, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Raj K. Ladher

raj-ladher@cdb.riken.jp

Affiliation Sensory Development, RIKEN Center for Developmental Biology, Chuo-ku, Kobe, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KO TK RKL. Performed the experiments: KO TK PO SSM MP. Analyzed the data: KO TK YS RKL. Contributed reagents/materials/analysis tools: SS MI NG KK. Wrote the paper: KO RKL.