Evolutionary Novelty in a Butterfly Wing Pattern through Enhancer Shuffling

• Richard W. R. Wallbank ,

  Contributed equally to this work with: Richard W. R. Wallbank, Simon W. Baxter

  Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom, Smithsonian Tropical Research Institution, Balboa, Ancón, Panama

  ⨯
• Simon W. Baxter ,

  Contributed equally to this work with: Richard W. R. Wallbank, Simon W. Baxter

  Affiliation School of Biological Sciences, University of Adelaide, Adelaide, Australia

  ⨯
• Carolina Pardo-Diaz,

  Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom, Smithsonian Tropical Research Institution, Balboa, Ancón, Panama, Biology Program, Faculty of Natural Sciences and Mathematics, Universidad del Rosario, Bogotá, D.C., Colombia

  ⨯
• Joseph J. Hanly,

  Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom, Smithsonian Tropical Research Institution, Balboa, Ancón, Panama

  ⨯
• Simon H. Martin,

  Affiliation Department of Zoology, University of Cambridge, Cambridge, United Kingdom

  ⨯
• James Mallet,

  Affiliations Smithsonian Tropical Research Institution, Balboa, Ancón, Panama, Organismic and Evolutionary Biology, Harvard University, Harvard, Massachusetts, United States of America

  ⨯
• Kanchon K. Dasmahapatra,

  Affiliations Smithsonian Tropical Research Institution, Balboa, Ancón, Panama, Department of Biology, University of York, York, United Kingdom

  ⨯
• Camilo Salazar,

  Affiliations Smithsonian Tropical Research Institution, Balboa, Ancón, Panama, Biology Program, Faculty of Natural Sciences and Mathematics, Universidad del Rosario, Bogotá, D.C., Colombia

  ⨯
• Mathieu Joron,

  Affiliations Smithsonian Tropical Research Institution, Balboa, Ancón, Panama, Institut de Systématique Evolution et Biodiversité, UMR 7205, CNRS MNHN UPMC EPHE, Muséum National d'Histoire Naturelle, CP50, Paris, France, Centre d’Ecologie Fonctionnelle et Evolutive, UMR 5175, CNRS–Université de Montpellier–Université Paul-Valéry–EPHE, Montpellier, France

  ⨯
• Nicola Nadeau,

  Affiliation Dept. of Animal and Plant Sciences, University of Sheffield, Sheffield, United Kingdom

  ⨯
• W. Owen McMillan,

  Affiliation Smithsonian Tropical Research Institution, Balboa, Ancón, Panama

  ⨯
• Chris D. Jiggins

  * E-mail: c.jiggins@zoo.cam.ac.uk

  Affiliations Department of Zoology, University of Cambridge, Cambridge, United Kingdom, Smithsonian Tropical Research Institution, Balboa, Ancón, Panama

  ⨯