About the Authors

Hiroko Sano

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

sano_hiroko@kurume-u.ac.jp

Affiliation Department of Molecular Genetics, Institute of Life Science, Kurume University, Kurume, Fukuoka, Japan

Akira Nakamura

Roles Funding acquisition, Investigation, Resources, Supervision, Writing – review & editing

Affiliation Department of Germline Development, Institute of Molecular Embryology and Genetics, and Graduate School of Pharmaceutical Sciences, Kumamoto University, Kumamoto, Kumamoto, Japan

Mariko Yamane

Roles Formal analysis, Writing – review & editing

Affiliation Department of Pluripotent Stem Cell Biology, Institute of Molecular Embryology and Genetics, and Graduate School of Medical Sciences, Kumamoto University, Kumamoto, Kumamoto, Japan

Hitoshi Niwa

Roles Formal analysis, Writing – review & editing

Affiliation Department of Pluripotent Stem Cell Biology, Institute of Molecular Embryology and Genetics, and Graduate School of Medical Sciences, Kumamoto University, Kumamoto, Kumamoto, Japan

Takashi Nishimura

Roles Investigation, Writing – review & editing

Affiliation Laboratory of Metabolic Regulation and Genetics, Institute for Molecular and Cellular Regulation, Gunma University, Maebashi, Gunma, Japan

Kimi Araki

Roles Investigation, Writing – review & editing

Affiliations Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Kumamoto, Japan, Center for Metabolic Regulation of Healthy Aging, Kumamoto University, Kumamoto, Kumamoto, Japan

Kazumasa Takemoto

Roles Investigation, Writing – review & editing

Affiliations Institute of Resource Development and Analysis, Kumamoto University, Kumamoto, Kumamoto, Japan, Department of Chromosome Biology, Institute of Molecular Embryology and Genetics, Kumamoto University, Kumamoto, Kumamoto, Japan

Kei-ichiro Ishiguro

Roles Investigation, Writing – review & editing

Affiliation Department of Chromosome Biology, Institute of Molecular Embryology and Genetics, Kumamoto University, Kumamoto, Kumamoto, Japan

Hiroki Aoki

Roles Investigation, Writing – review & editing

Affiliation Cardiovascular Research Institute, Kurume University, Kurume, Fukuoka, Japan

Yuzuru Kato

Roles Software, Writing – review & editing

Affiliations Mammalian Development Laboratory, Department of Gene Function and Phenomics, National Institute of Genetics, Mishima, Shizuoka, Japan, Department of Genetics, SOKENDAI, Mishima, Shizuoka, Japan

Masayasu Kojima

Roles Funding acquisition, Investigation, Writing – review & editing

Affiliation Department of Molecular Genetics, Institute of Life Science, Kurume University, Kurume, Fukuoka, Japan

Competing Interests

The authors have declared that no competing interests exist.