About the Authors

Dan Fu

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology, Validation

Affiliation State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China

Guangshun Zhang

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology, Software, Validation, Visualization

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China, College of Life Science, Nankai University, Tianjin, China

Yuhui Wang

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China

Zheng Zhang

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology

Affiliation State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China

Hengrui Hu

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliations Center for Biosafety Mega-Science, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China, Guangzhou Institutes of Biomedicine and Health, Chinese Academy of Sciences, Guangzhou, China

Shu Shen

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology

Affiliations Center for Biosafety Mega-Science, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China, State Key Laboratory of Virology and National Virus Resource Center, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China

Jun Wu

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Data curation, Investigation, Methodology, Software, Visualization, Writing – original draft

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Bo Li

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China

Xin Li

Contributed equally to this work with: Dan Fu, Guangshun Zhang, Yuhui Wang, Zheng Zhang, Hengrui Hu, Shu Shen, Jun Wu, Bo Li, Xin Li

Roles Investigation, Methodology, Software, Visualization

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China, Shanghai Institute for Advanced Immunochemical Studies, ShanghaiTech University, Shanghai, China

Yaohui Fang

Roles Investigation, Methodology

Affiliation State Key Laboratory of Virology and National Virus Resource Center, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China

Jia Liu

Roles Investigation, Methodology

Affiliation Center for Biosafety Mega-Science, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China

Qiao Wang

Roles Investigation, Methodology, Resources, Supervision

Affiliation Key Laboratory of Medical Molecular Virology (MOE/NHC/CAMS), School of Basic Medical Sciences, Shanghai Medical College, Fudan University, Shanghai, China

Yunjiao Zhou

Roles Investigation, Methodology

Affiliation Key Laboratory of Medical Molecular Virology (MOE/NHC/CAMS), School of Basic Medical Sciences, Shanghai Medical College, Fudan University, Shanghai, China

Wei Wang

Roles Investigation, Methodology

Affiliation Shanghai Institute for Advanced Immunochemical Studies, ShanghaiTech University, Shanghai, China

Yufeng Li

Roles Methodology, Project administration

Affiliation Center for Biosafety Mega-Science, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China

Zhonghua Lu

Roles Investigation, Methodology, Software

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Xiaoxiao Wang

Roles Investigation, Methodology, Supervision

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Cui Nie

Roles Investigation, Methodology

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Yujie Tian

Roles Investigation, Methodology

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, Frontiers Science Center for Cell Responses, Nankai University, Tianjin, China

Da Chen

Roles Methodology, Project administration, Software, Visualization

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China, Frontiers Science Center for Cell Responses, Nankai University, Tianjin, China

Yuan Wang

Roles Investigation, Methodology, Supervision

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China, Frontiers Science Center for Cell Responses, Nankai University, Tianjin, China

Xingdong Zhou

Roles Investigation, Methodology

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China, Frontiers Science Center for Cell Responses, Nankai University, Tianjin, China

Qisheng Wang

Roles Software, Supervision, Validation, Visualization

Affiliation Shanghai Synchrotron Radiation Facility, Shanghai Advanced Research Institute, Chinese Academy of Sciences, Shanghai, China

Feng Yu

Roles Investigation, Methodology, Software

Affiliation Shanghai Synchrotron Radiation Facility, Shanghai Advanced Research Institute, Chinese Academy of Sciences, Shanghai, China

Chen Zhang

Roles Investigation, Methodology

Affiliation State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China

Changjing Deng

Roles Investigation, Methodology

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Liang Zhou

Roles Investigation, Methodology

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Guangkuo Guan

Roles Investigation, Methodology

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Na Shao

Roles Investigation, Methodology

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Zhiyong Lou

Roles Conceptualization, Project administration, Resources

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliation MOE Key Laboratory of Protein Science & Collaborative Innovation Center of Biotherapy, School of Medicine, Tsinghua University, Beijing, China

Fei Deng

Roles Investigation, Methodology, Project administration, Supervision

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliations Center for Biosafety Mega-Science, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China, State Key Laboratory of Virology and National Virus Resource Center, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China

Hongkai Zhang

Roles Conceptualization, Funding acquisition, Supervision, Writing – original draft

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, Shanghai Institute for Advanced Immunochemical Studies, ShanghaiTech University, Shanghai, China, Frontiers Science Center for Cell Responses, Nankai University, Tianjin, China

Xinwen Chen

Roles Conceptualization, Investigation, Resources, Writing – original draft

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliations Guangzhou Institutes of Biomedicine and Health, Chinese Academy of Sciences, Guangzhou, China, Guangzhou Laboratory, Guangzhou International Bio-Island, Guangzhou, Guangdong, China

Manli Wang

Roles Investigation, Methodology, Supervision, Writing – review & editing

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliation Center for Biosafety Mega-Science, Wuhan Institute of Virology, Chinese Academy of Sciences, Wuhan, Hubei, China

Louis Liu

Roles Conceptualization, Funding acquisition, Investigation, Methodology

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliation Harbour Biomed (Suzhou) Co. Ltd., Suzhou Industrial Park, Suzhou, China

Zihe Rao

Roles Conceptualization, Funding acquisition, Project administration, Supervision, Writing – original draft

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, College of Life Science, Nankai University, Tianjin, China, Shanghai Institute for Advanced Immunochemical Studies, ShanghaiTech University, Shanghai, China

Yu Guo

Roles Conceptualization, Formal analysis, Funding acquisition, Project administration, Supervision, Writing – original draft, Writing – review & editing

louzy@mail.tsinghua.edu.cn (ZL); df@wh.iov.cn (FD); hongkai@nankai.edu.cn (HZ); chen_xinwen@gibh.ac.cn (XC); wangml@wh.iov.cn (MW); Louis.Liu@harbourbiomed.com (LL); raozh@mail.tsinghua.edu.cn (ZR); guoyu@nankai.edu.cn (YG)

Affiliations State Key Laboratory of Medicinal Chemical Biology and College of Pharmacy, Nankai University, Tianjin, China, Frontiers Science Center for Cell Responses, Nankai University, Tianjin, China, Guangzhou Laboratory, Guangzhou International Bio-Island, Guangzhou, Guangdong, China

Competing Interests

I have read the journal’s policy and the authors of this manuscript have the following competing interests: Y.G., Z.R., L.L., J.W., Z.L., D.F., X.W. and C.N. are inventors in a pending patent application filed on the reported antibodies. L.L., J.W., Z.L., X.W., C.N., C.D., L.Z., G.G. and N.S. are employees of Harbour Biomed (Suzhou) Co. Ltd. Other authors declare no competing interests. All reagents and information presented in this study are available from corresponding authors upon reasonable request.