About the Authors

Lizhen Wu

Contributed equally to this work with: Lizhen Wu, Jian Cao

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Jian Cao

Contributed equally to this work with: Lizhen Wu, Jian Cao

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Wesley L. Cai

Roles Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Visualization, Writing – review & editing

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Sabine M. Lang

Roles Formal analysis, Investigation, Methodology, Project administration, Validation, Visualization

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

John R. Horton

Roles Data curation, Formal analysis, Investigation, Methodology, Validation, Visualization

Affiliation Department of Molecular and Cellular Oncology, The University of Texas MD Anderson Cancer Center, Houston, Texas, United States of America

Daniel J. Jansen

Roles Investigation, Methodology, Resources, Validation, Visualization, Writing – review & editing

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Zongzhi Z. Liu

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Jocelyn F. Chen

Roles Formal analysis, Investigation, Validation

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Meiling Zhang

Roles Investigation, Validation

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Bryan T. Mott

Roles Investigation, Methodology, Resources, Validation, Visualization

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Katherine Pohida

Roles Investigation, Methodology, Resources, Validation, Visualization

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Ganesha Rai

Roles Investigation, Methodology, Resources, Validation, Visualization

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Stephen C. Kales

Roles Investigation, Methodology, Resources, Validation, Visualization, Writing – review & editing

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Mark J. Henderson

Roles Investigation, Methodology, Resources, Validation, Visualization, Writing – review & editing

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Xin Hu

Roles Investigation, Methodology, Resources, Validation, Visualization

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Ajit Jadhav

Roles Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

David J. Maloney

Roles Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Anton Simeonov

Roles Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – review & editing

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Shu Zhu

Roles Methodology, Resources, Writing – review & editing

Affiliation Institute of Immunology and the CAS Key Laboratory of Innate Immunity and Chronic Disease, School of Life Sciences and Medical Center, University of Science and Technology of China, Hefei, China

Akiko Iwasaki

Roles Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Visualization, Writing – review & editing

Affiliations Department of Immunobiology, Yale School of Medicine, New Haven, Connecticut, United States of America, Howard Hughes Medical Institute, Chevy Chase, Maryland, United States of America

Matthew D. Hall

Roles Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – review & editing

Affiliation National Center for Advancing Translational Sciences, National Institutes of Health, Rockville, Maryland, United States of America

Xiaodong Cheng

Roles Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Molecular and Cellular Oncology, The University of Texas MD Anderson Cancer Center, Houston, Texas, United States of America

Gerald S. Shadel

Roles Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – review & editing

Affiliations Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America, Department of Genetics, Yale School of Medicine, New Haven, Connecticut, United States of America, Salk Institute for Biological Studies, La Jolla, California, United States of America

Qin Yan

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Resources, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

qin.yan@yale.edu

Affiliation Department of Pathology, Yale School of Medicine, New Haven, Connecticut, United States of America

Competing Interests

The authors have declared that no competing interests exist.