About the Authors

Roger B. J. Benson

roger.benson@earth.ox.ac.uk

Affiliation Department of Earth Sciences, University of Oxford, Oxford, United Kingdom

Richard J. Butler

Affiliation School of Geography, Earth and Environmental Sciences, University of Birmingham, Birmingham, United Kingdom

John Alroy

Affiliation Department of Biological Sciences, Macquarie University, New South Wales, Australia

Philip D. Mannion

Affiliation Department of Earth Science and Engineering, Imperial College London, London, United Kingdom

Matthew T. Carrano

Affiliation Department of Paleobiology, Smithsonian Institution, Washington, D.C., United States of America

Graeme T. Lloyd

Affiliation Department of Biological Sciences, Macquarie University, New South Wales, Australia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RBJB JA RJB PDM GTL. Analyzed the data: RBJB. Contributed reagents/materials/analysis tools: MTC JA RJB PDM RBJB. Wrote the paper: RBJB RJB PDM JA MTC GTL.