About the Authors

Catherine Hobaiter

Affiliations School of Psychology and Neuroscience, University of St. Andrews, Fife, United Kingdom, Budongo Conservation Field Station, Masindi, Uganda

Timothée Poisot

Affiliations Département de Biologie, Chimie et Géographie, Université du Québec à Rimouski, Rimouski, Québec, Canada, Québec Centre for Biodiversity Sciences, Montréal, Québec, Canada

Klaus Zuberbühler

Affiliations School of Psychology and Neuroscience, University of St. Andrews, Fife, United Kingdom, Budongo Conservation Field Station, Masindi, Uganda, Department of Comparative Cognition, Institute of Biology, University of Neuchâtel, Neuchâtel, Switzerland

William Hoppitt

Affiliation Animal and Environment Research Group, Anglia Ruskin University, Cambridge, United Kingdom

Thibaud Gruber

thibaud.gruber@gmail.com

Affiliations Budongo Conservation Field Station, Masindi, Uganda, Department of Comparative Cognition, Institute of Biology, University of Neuchâtel, Neuchâtel, Switzerland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: CH. Performed the experiments: CH. Analyzed the data: CH TG WH TP. Contributed reagents/materials/analysis tools: TP WH. Wrote the paper: TG CH KZ WH TP.