About the Authors

Adam D. Hayward

adam.hayward@ed.ac.uk (ADH); algraham@princeton.edu (ALG)

Affiliation Department of Animal and Plant Sciences, University of Sheffield, Sheffield, United Kingdom

Daniel H. Nussey

Affiliations Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom, Institute of Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom, Centre for Immunity, Infection and Evolution, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Alastair J. Wilson

Affiliation Centre for Ecology and Conservation, University of Exeter, Cornwall Campus, Penryn, Cornwall, United Kingdom

Camillo Berenos

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Jill G. Pilkington

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Kathryn A. Watt

Affiliations Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom, Institute of Immunology and Infection Research, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom, Centre for Immunity, Infection and Evolution, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Josephine M. Pemberton

Affiliation Institute of Evolutionary Biology, School of Biological Sciences, University of Edinburgh, Edinburgh, United Kingdom

Andrea L. Graham

adam.hayward@ed.ac.uk (ADH); algraham@princeton.edu (ALG)

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: ADH DHN ALG. Analyzed the data: ADH DHN AJW. Contributed reagents/materials/analysis tools: JMP CB. Contributed to the writing of the manuscript: ADH DHN AJW CB KAW JMP ALG. Conducted fieldwork: JGP.