About the Authors

Estienne C. Swart

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

John R. Bracht

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Vincent Magrini

Affiliations The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Patrick Minx

Affiliation The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America

Xiao Chen

Affiliation Department of Molecular Biology, Princeton University, Princeton, New Jersey, United States of America

Yi Zhou

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Jaspreet S. Khurana

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Aaron D. Goldman

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Mariusz Nowacki

Affiliations Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America, Institute of Cell Biology, University of Bern, Bern, Switzerland

Klaas Schotanus

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Seolkyoung Jung

Affiliation Janelia Farm Research Campus, Howard Hughes Medical Institute, Ashburn, Virginia, United States of America

Robert S. Fulton

Affiliations The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Amy Ly

Affiliation The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America

Sean McGrath

Affiliation The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America

Kevin Haub

Affiliation The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America

Jessica L. Wiggins

Affiliation Sequencing Core Facility, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Donna Storton

Affiliation Sequencing Core Facility, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

John C. Matese

Affiliation Sequencing Core Facility, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Lance Parsons

Affiliation Bioinformatics Group, Lewis-Sigler Institute for Integrative Genomics, Princeton University, Princeton, New Jersey, United States of America

Wei-Jen Chang

Affiliation Department of Biology, Hamilton College, Clinton, New York, United States of America

Michael S. Bowen

Affiliation Biology Department, Bradley University, Peoria, Illinois, United States of America

Nicholas A. Stover

Affiliation Biology Department, Bradley University, Peoria, Illinois, United States of America

Thomas A. Jones

Affiliation Janelia Farm Research Campus, Howard Hughes Medical Institute, Ashburn, Virginia, United States of America

Sean R. Eddy

Affiliation Janelia Farm Research Campus, Howard Hughes Medical Institute, Ashburn, Virginia, United States of America

Glenn A. Herrick

Affiliation Biology Department, University of Utah, Salt Lake City, Utah, United States of America

Thomas G. Doak

Affiliation Department of Biology, University of Indiana, Bloomington, Indiana, United States of America

Richard K. Wilson

Affiliations The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Elaine R. Mardis

Affiliations The Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America, Department of Genetics, Washington University School of Medicine, St. Louis, Missouri, United States of America

Laura F. Landweber

lfl@princeton.edu

Affiliation Department of Ecology and Evolutionary Biology, Princeton University, Princeton, New Jersey, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Designed the software used in analysis: ECS. Conceived and designed the experiments: ECS VM YZ TGD JRB RKW ERM LFL. Performed the experiments: ECS JRB VM TGD JSK MN KS AL SM KH JLB DS. Analyzed the data: ECS VM PM XC YZ ADG SJ WC RSF. Contributed reagents/materials/analysis tools: ECS JRB MN KS VM AL SM KH JCM LP TGD SJ TAJ SRE MSB NAS. Wrote the paper: ECS JRB VM XC TGD GAH LFL.