About the Authors

Vivian G. Cheung

vcheung@mail.med.upenn.edu

Affiliations Howard Hughes Medical Institute, Philadelphia, Pennsylvania, United States of America, Department of Pediatrics, University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America, Department of Genetics, University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America, University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Renuka R. Nayak

Affiliation Medical Scientist Training Program, University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Isabel Xiaorong Wang

Affiliation Howard Hughes Medical Institute, Philadelphia, Pennsylvania, United States of America

Susannah Elwyn

Affiliation University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Sarah M. Cousins

Affiliation University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Michael Morley

Affiliation University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Richard S. Spielman

† Deceased.

Affiliation Department of Genetics, University of Pennsylvania, The Children's Hospital of Philadelphia, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

The author(s) have made the following declarations about their contributions: Conceived and designed the experiments: VGC RSS. Performed the experiments: RRN IXW SE. Analyzed the data: VGC RRN IXW SE SMC MM RSS. Wrote the paper: VGC RRN IXW RSS.