About the Authors

Jason B Wolf

To whom correspondence should be addressed. E-mail: Jason@evolutionarygenetics.org

Affiliation Faculty of Life Sciences, The University of Manchester, Manchester, United Kingdom

Reinmar Hager

Affiliation Faculty of Life Sciences, The University of Manchester, Manchester, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

JBW and RH conceived the idea for the model. JW derived the model. JBW and RH wrote the paper.