About the Authors

David L Reed

To whom correspondence should be addressed. E-mail: dreed@flmnh.ufl.edu

Affiliation Florida Museum of Natural History, Dickinson Hall, University of Florida, Gainesville, Florida, United States of America

Vincent S Smith

Affiliation Graham Kerr Building, DEEB, IBLS, University of Glasgow, Glasgow, Scotland

Shaless L Hammond

Affiliation Department of Biology, University of Utah, Salt Lake City, Utah, United States of America

Alan R Rogers

Affiliation Department of Anthropology, University of Utah, Salt Lake City, Utah, United States of America

Dale H Clayton

Affiliation Department of Biology, University of Utah, Salt Lake City, Utah, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

DLR, VSS, and DHC designed the study. DLR, VSS, and SLH collected and analyzed the data. DLR, VSS, ARR, and DHC wrote the paper.