About the Authors

Elizabeth A Hadly

To whom correspondence should be addressed. E-mail: hadly@stanford.edu

Affiliation Department of Biological Sciences, Stanford University, Stanford, California, United States of America

Uma Ramakrishnan

Contributed equally to this work with: Uma Ramakrishnan, Yvonne L Chan, Marcel van Tuinen

Affiliation Department of Biological Sciences, Stanford University, Stanford, California, United States of America

Yvonne L Chan

Contributed equally to this work with: Uma Ramakrishnan, Yvonne L Chan, Marcel van Tuinen

Affiliation Department of Biological Sciences, Stanford University, Stanford, California, United States of America

Marcel van Tuinen

Contributed equally to this work with: Uma Ramakrishnan, Yvonne L Chan, Marcel van Tuinen

Affiliation Department of Biological Sciences, Stanford University, Stanford, California, United States of America

Kim O'Keefe

Affiliation Department of Biological Sciences, Stanford University, Stanford, California, United States of America

Paula A Spaeth

Affiliation Department of Biological Sciences, Stanford University, Stanford, California, United States of America

Chris J Conroy

Affiliation Museum of Vertebrate Zoology, University of California, Berkeley, Berkeley, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

EAH conceived and designed the experiments. YLC, MV, KOK, PAS, and CJC performed the experiments. EAH, UR, YLC, and MV analyzed the data. EAH and UR contributed reagents/materials/analysis tools. EAH, UR, YLC, MV, and KOK wrote the paper.