About the Authors

Jun R Huh

Affiliation Division of Biology, California Institute of Technology, Pasadena, California, United States of America

Stephanie Y Vernooy

Affiliation Division of Biology, California Institute of Technology, Pasadena, California, United States of America

Hong Yu

Affiliation Division of Biology, California Institute of Technology, Pasadena, California, United States of America

Nieng Yan

Affiliation Department of Molecular Biology, Lewis Thomas Laboratory, Princeton University, Princeton, New Jersey, United States of America

Yigong Shi

Affiliation Department of Molecular Biology, Lewis Thomas Laboratory, Princeton University, Princeton, New Jersey, United States of America

Ming Guo

Affiliation Department of Neurology, Brain Research Institute, The David Geffen School of Medicine at the University of California at Los Angeles, Los Angeles, California, United States of America

Bruce A Hay

Affiliation Division of Biology, California Institute of Technology, Pasadena, California, United States of America

Competing Interests

The authors have declared that no conflicts of interest exist.

Author Contributions

JRH and BAH conceived and designed the experiments. JRH and SYV performed the experiments. JRH and BAH analyzed the data. JRH, HY, NY, YS, MG, and BAH contributed reagents/materials/analysis tools. JRH and BAH wrote the paper.