Quantifying Missing Heritability at Known GWAS Loci

• Alexander Gusev ,

  * E-mail: agusev@hsph.harvard.edu (AG); aprice@hsph.harvard.edu (ALP)

  Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America

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• Gaurav Bhatia,

  Affiliations Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America, Harvard-Massachusetts Institute of Technology (MIT) Division of Health, Science and Technology, Cambridge, Massachusetts, United States of America

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• Noah Zaitlen,

  Affiliation Department of Medicine Lung Biology Center, University of California San Francisco, San Francisco, California, United States of America

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• Bjarni J. Vilhjalmsson,

  Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America

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• Dorothée Diogo,

  Affiliations Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America, Division of Rheumatology, Immunology, and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

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• Eli A. Stahl,

  Affiliations Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America, Division of Rheumatology, Immunology, and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

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• Peter K. Gregersen,

  Affiliation The Feinstein Institute for Medical Research, North Shore-Long Island Jewish Health System, Manhasset, New York, United States of America

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• Jane Worthington,

  Affiliation Arthritis Research UK Epidemiology Unit, University of Manchester, Manchester Academic Health Sciences Centre, Manchester, United Kingdom

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• Lars Klareskog,

  Affiliation Rheumatology Unit, Department of Medicine, Karolinska Institutet and Karolinska University Hospital Solna, Stockholm, Sweden

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• Soumya Raychaudhuri,

  Affiliations Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America, Division of Rheumatology, Immunology, and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Arthritis Research UK Epidemiology Unit, University of Manchester, Manchester Academic Health Sciences Centre, Manchester, United Kingdom

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• Robert M. Plenge,

  Affiliations Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America, Division of Rheumatology, Immunology, and Allergy, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America, Division of Genetics, Brigham and Women's Hospital, Harvard Medical School, Boston, Massachusetts, United States of America

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• Bogdan Pasaniuc,

  Affiliation Department of Pathology and Laboratory Medicine, Geffen School of Medicine at UCLA, Los Angeles, California, United States of America

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• Alkes L. Price

  * E-mail: agusev@hsph.harvard.edu (AG); aprice@hsph.harvard.edu (ALP)

  Affiliations Department of Epidemiology, Harvard School of Public Health, Boston, Massachusetts, United States of America, Department of Biostatistics, Harvard School of Public Health, Boston, Massachusetts, United States of America, Medical and Population Genetics Program, Broad Institute, Cambridge, Massachusetts, United States of America

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