About the Authors

Cassandra N. Spracklen

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America, Department of Biostatistics and Epidemiology, University of Massachusetts, Amherst, Massachusetts, United States of America

Apoorva K. Iyengar

Roles Investigation, Validation, Writing – original draft, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Swarooparani Vadlamudi

Roles Investigation, Validation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Chelsea K. Raulerson

Roles Writing – original draft, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Anne U. Jackson

Roles Data curation, Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Sarah M. Brotman

Roles Formal analysis, Investigation, Validation, Writing – original draft, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Ying Wu

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Maren E. Cannon

Roles Investigation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

James P. Davis

Roles Investigation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Aaron T. Crain

Roles Investigation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Kevin W. Currin

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Hannah J. Perrin

Roles Formal analysis, Investigation, Writing – review & editing

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Narisu Narisu

Roles Resources, Writing – review & editing

Affiliation National Human Genome Research Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Heather M. Stringham

Roles Data curation, Resources, Writing – review & editing

Affiliation Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Christian Fuchsberger

Roles Resources, Writing – review & editing

Affiliations Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America, Center for Biomedicine, European Academy of Bolzano/Bozen, University of Lübeck, Bolzano/Bozen, Italy

Adam E. Locke

Roles Resources, Writing – review & editing

Affiliations Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America, McDonnell Genome Institute, Washington University School of Medicine, St. Louis, Missouri, United States of America

Ryan P. Welch

Roles Resources, Writing – review & editing

Affiliation Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Johanna K. Kuusisto

Roles Supervision, Writing – review & editing

Affiliation Department of Medicine, University of Eastern Finland and Kuopio University Hospital, Kuopio, Finland

Päivi Pajukanta

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation Department of Human Genetics, University of California, Los Angeles, California, United States of America

Laura J. Scott

Roles Resources, Supervision, Writing – review & editing

Affiliation Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Yun Li

Roles Resources, Supervision, Writing – review & editing

Affiliations Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America, Department of Biostatistics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Francis S. Collins

Roles Conceptualization, Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliation National Human Genome Research Institute, National Institutes of Health, Bethesda, Maryland, United States of America

Michael Boehnke

Roles Conceptualization, Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliation Department of Biostatistics and Center for Statistical Genetics, School of Public Health, University of Michigan, Ann Arbor, Michigan, United States of America

Markku Laakso

Roles Conceptualization, Data curation, Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliation Department of Medicine, University of Eastern Finland and Kuopio University Hospital, Kuopio, Finland

Karen L. Mohlke

Roles Conceptualization, Funding acquisition, Resources, Supervision, Validation, Writing – original draft, Writing – review & editing

mohlke@med.unc.edu

Affiliation Department of Genetics, University of North Carolina, Chapel Hill, North Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.