About the Authors

Mateus H. Gouveia

Contributed equally to this work with: Mateus H. Gouveia, Andrew W. Bergen, Victor Borda, Kelly Nunes

Roles Data curation, Formal analysis, Investigation, Methodology, Software, Supervision, Writing – original draft, Writing – review & editing

Affiliations Instituto de Pesquisa René Rachou, Fundação Oswaldo Cruz, Belo Horizonte, Minas Gerais, Brazil, Departamento de Biologia Geral, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais, Belo Horizonte, Minas Gerais, Brazil, Center for Research on Genomics & Global Health, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Andrew W. Bergen

Contributed equally to this work with: Mateus H. Gouveia, Andrew W. Bergen, Victor Borda, Kelly Nunes

Roles Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Current address: Oregon Research Institute, Eugene, Oregon

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Victor Borda

Contributed equally to this work with: Mateus H. Gouveia, Andrew W. Bergen, Victor Borda, Kelly Nunes

Roles Data curation, Formal analysis, Investigation, Methodology, Software, Writing – original draft, Writing – review & editing

Affiliation Departamento de Biologia Geral, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais, Belo Horizonte, Minas Gerais, Brazil

Kelly Nunes

Contributed equally to this work with: Mateus H. Gouveia, Andrew W. Bergen, Victor Borda, Kelly Nunes

Roles Formal analysis, Investigation, Methodology, Software, Writing – original draft, Writing – review & editing

Affiliation Departamento de Genética e Biologia Evolutiva, Instituto de Biociências, Universidade de São Paulo, São Paulo, Brazil

Thiago P. Leal

Roles Formal analysis, Investigation, Methodology, Software, Writing – original draft

Affiliations Departamento de Biologia Geral, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais, Belo Horizonte, Minas Gerais, Brazil, Department of Statistics, Universidade Federal de Minas Gerais, Belo Horizonte, Minas Gerais, Brazil

Martin D. Ogwang

Roles Methodology, Supervision

Affiliation EMBLEM Study, African Field Epidemiology Network, Kampala, Uganda

Edward D. Yeboah

Roles Methodology

Affiliation University of Ghana Medical School, Accra, Ghana

James E. Mensah

Roles Methodology

Affiliation University of Ghana Medical School, Accra, Ghana

Tobias Kinyera

Roles Methodology

Affiliation EMBLEM Study, African Field Epidemiology Network, Kampala, Uganda

Isaac Otim

Roles Methodology

Affiliation EMBLEM Study, African Field Epidemiology Network, Kampala, Uganda

Hadijah Nabalende

Roles Methodology

Affiliation EMBLEM Study, African Field Epidemiology Network, Kampala, Uganda

Ismail D. Legason

Roles Methodology

Affiliation EMBLEM Study, African Field Epidemiology Network, Kampala, Uganda

Sununguko Wata Mpoloka

Roles Methodology

Affiliation Department of Biological Sciences, University of Botswana, Gaborone, Botswana

Gaonyadiwe George Mokone

Roles Methodology

Affiliation Department of Biomedical Sciences, University of Botswana School of Medicine, Gaborone, Botswana

Patrick Kerchan

Roles Methodology

Affiliation EMBLEM Study, African Field Epidemiology Network, Kampala, Uganda

Kishor Bhatia

Roles Methodology

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Steven J. Reynolds

Roles Methodology

Affiliation Division of Intramural Research, National Institute of Allergy and Infectious Diseases, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Richard B. Birtwum

Roles Methodology

Affiliation University of Ghana Medical School, Accra, Ghana

Andrew A. Adjei

Roles Methodology

Affiliation University of Ghana Medical School, Accra, Ghana

Yao Tettey

Roles Methodology

Affiliation University of Ghana Medical School, Accra, Ghana

Evelyn Tay

Roles Methodology

Affiliation University of Ghana Medical School, Accra, Ghana

Robert Hoover

Roles Methodology

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Ruth M. Pfeiffer

Roles Methodology

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Robert J. Biggar

Roles Methodology

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

James J. Goedert

Roles Methodology

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Ludmila Prokunina-Olsson

Roles Methodology, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Translational Genomics, Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Michael Dean

Roles Methodology

Affiliation Laboratory of Translational Genomics, Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Meredith Yeager

Roles Methodology

Affiliation Cancer Genomics Research Laboratory, Leidos Biomedical Research, Frederick National Laboratory for Cancer Research, US Department of Health and Human Services, Frederick, Maryland, United States of America

M. Fernanda Lima-Costa

Roles Methodology, Writing – review & editing

Affiliation Instituto de Pesquisa René Rachou, Fundação Oswaldo Cruz, Belo Horizonte, Minas Gerais, Brazil

Ann W. Hsing

Roles Methodology

Affiliation Stanford Cancer Institute, Stanford University, Stanford, California, United States of America

Sarah A. Tishkoff

Roles Methodology, Writing – review & editing

Affiliation Department of Genetics and Biology, University of Pennsylvania, Philadelphia, United States of America

Stephen J. Chanock

Roles Writing – review & editing

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Eduardo Tarazona-Santos

Roles Writing – original draft, Writing – review & editing

‡ These authors also contributed equally to this work.

Affiliation Departamento de Biologia Geral, Instituto de Ciências Biológicas, Universidade Federal de Minas Gerais, Belo Horizonte, Minas Gerais, Brazil

Sam M. Mbulaiteye

Roles Conceptualization, Data curation, Investigation, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

mbulaits@mail.nih.gov

‡ These authors also contributed equally to this work.

Affiliation Division of Cancer Epidemiology and Genetics, National Cancer Institute, National Institutes of Health, US Department of Health and Human Services, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.