About the Authors

Ou Fang

Contributed equally to this work with: Ou Fang, Xiaohua Hu

Roles Conceptualization, Methodology, Writing – original draft, Writing – review & editing

Affiliations Laboratory of Population and Quantitative Genetics, Institute of Biostatistics and Genetics, School of Life Sciences, Fudan University, Shanghai, China, School of Biosciences, the University of Birmingham, Birmingham, United Kingdom, Department of Evolution and Ecology, School of Life Sciences, Fudan University, Shanghai, China

Xiaohua Hu

Contributed equally to this work with: Ou Fang, Xiaohua Hu

Roles Conceptualization, Methodology, Writing – original draft, Writing – review & editing

Affiliation Laboratory of Population and Quantitative Genetics, Institute of Biostatistics and Genetics, School of Life Sciences, Fudan University, Shanghai, China

Lin Wang

Roles Methodology

Affiliation Laboratory of Population and Quantitative Genetics, Institute of Biostatistics and Genetics, School of Life Sciences, Fudan University, Shanghai, China

Ning Jiang

Roles Data curation, Software

Affiliation Laboratory of Population and Quantitative Genetics, Institute of Biostatistics and Genetics, School of Life Sciences, Fudan University, Shanghai, China

Jixuan Yang

Roles Methodology

Affiliation Laboratory of Population and Quantitative Genetics, Institute of Biostatistics and Genetics, School of Life Sciences, Fudan University, Shanghai, China

Bo Li

Roles Writing – original draft

Affiliation Department of Evolution and Ecology, School of Life Sciences, Fudan University, Shanghai, China

Zewei Luo

Roles Conceptualization, Writing – original draft, Writing – review & editing

z.luo@bham.ac.uk, zwluo@fudan.edu.cn

Affiliations Laboratory of Population and Quantitative Genetics, Institute of Biostatistics and Genetics, School of Life Sciences, Fudan University, Shanghai, China, School of Biosciences, the University of Birmingham, Birmingham, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.