About the Authors

James C. Hart

Roles Conceptualization, Formal analysis, Investigation, Writing – original draft, Writing – review & editing

Affiliation Department of Molecular and Cell Biology, University of California-Berkeley, CA, United States of America

Nicholas A. Ellis

Roles Investigation, Writing – review & editing

Affiliation Department of Molecular and Cell Biology, University of California-Berkeley, CA, United States of America

Michael B. Eisen

Roles Conceptualization, Writing – review & editing

Affiliations Department of Molecular and Cell Biology, University of California-Berkeley, CA, United States of America, Howard Hughes Medical Institute, University of California, Berkeley, CA, United States of America

Craig T. Miller

Roles Conceptualization, Funding acquisition, Writing – original draft, Writing – review & editing

ctmiller@berkeley.edu

Affiliation Department of Molecular and Cell Biology, University of California-Berkeley, CA, United States of America

Competing Interests

I have read the journal's policy and the authors of this manuscript have the following competing interests: MBE is a cofounder and member of the Board of Directors of PLOS. This does not alter the authors' adherence to all the PLOS policies on sharing data and materials.