About the Authors

Seth D. Kasowitz

Contributed equally to this work with: Seth D. Kasowitz, Jun Ma

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliation Department of Biomedical Sciences, University of Pennsylvania, Philadelphia, United States of America

Jun Ma

Contributed equally to this work with: Seth D. Kasowitz, Jun Ma

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliations Department of Biomedical Sciences, University of Pennsylvania, Philadelphia, United States of America, Department of Biology, University of Pennsylvania, Philadelphia, United States of America

Stephen J. Anderson

Roles Data curation, Formal analysis, Investigation, Methodology, Writing – original draft, Writing – review & editing

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, United States of America

N. Adrian Leu

Roles Investigation, Methodology, Resources, Writing – review & editing

Affiliation Department of Biomedical Sciences, University of Pennsylvania, Philadelphia, United States of America

Yang Xu

Roles Investigation, Methodology, Writing – review & editing

Affiliation Department of Biomedical Sciences, University of Pennsylvania, Philadelphia, United States of America

Brian D. Gregory

Roles Conceptualization, Formal analysis, Funding acquisition, Resources, Supervision, Writing – review & editing

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, United States of America

Richard M. Schultz

Roles Conceptualization, Formal analysis, Funding acquisition, Supervision, Writing – review & editing

Affiliations Department of Biology, University of Pennsylvania, Philadelphia, United States of America, Department of Anatomy, Physiology and Cell Biology, School of Veterinary Medicine, University of California, Davis, Davis, United States of America

P. Jeremy Wang

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Resources, Supervision, Validation, Writing – original draft, Writing – review & editing

pwang@vet.upenn.edu

Affiliation Department of Biomedical Sciences, University of Pennsylvania, Philadelphia, United States of America

Competing Interests

The authors have declared that no competing interests exist.