About the Authors

Jia He

Roles Conceptualization, Formal analysis, Investigation, Methodology, Writing – original draft

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Mingli Xu

Roles Investigation

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Matthew R. Willmann

Roles Conceptualization, Formal analysis, Investigation

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Kevin McCormick

Roles Data curation, Software

Affiliation Department of Soil and Plant Sciences, University of Delaware, Newark, Delaware, United States of America

Tieqiang Hu

Roles Investigation

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Li Yang

Roles Investigation, Resources

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Colby G. Starker

Roles Methodology

Affiliation Department of Genetics, Cell Biology and Development and Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota, United States of America

Daniel F. Voytas

Roles Funding acquisition

Affiliation Department of Genetics, Cell Biology and Development and Center for Genome Engineering, University of Minnesota, Minneapolis, Minnesota, United States of America

Blake C. Meyers

Roles Funding acquisition, Project administration, Supervision

Affiliation Department of Soil and Plant Sciences, University of Delaware, Newark, Delaware, United States of America

R. Scott Poethig

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Project administration, Supervision, Writing – review & editing

spoethig@sas.upenn.edu

Affiliation Department of Biology, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.