About the Authors

Takeshi Yasuda

Contributed equally to this work with: Takeshi Yasuda, Wataru Kagawa

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Project administration, Resources, Validation, Visualization, Writing – original draft, Writing – review & editing

yasuda.takeshi@qst.go.jp (TY); tajima.katsushi@qst.go.jp (KT)

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Wataru Kagawa

Contributed equally to this work with: Takeshi Yasuda, Wataru Kagawa

Roles Formal analysis, Investigation, Resources, Validation, Visualization, Writing – review & editing

Affiliation Program in Chemistry and Life Science, Department of Interdisciplinary Science and Engineering, School of Science and Engineering, Meisei University, Hodokubo, Hino-shi, Tokyo, Japan

Tomoo Ogi

Roles Investigation, Resources, Validation, Visualization, Writing – review & editing

Affiliation Department of Genetics, Research Institute of Environmental Medicine, Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Japan

Takamitsu A. Kato

Roles Formal analysis, Investigation

Affiliation Department of Environmental and Radiological Health Sciences, Colorado State University, Fort Collins, CO, United States of America

Takehiro Suzuki

Roles Formal analysis, Investigation

Affiliation Biomolecular Characterization Unit, RIKEN Center for Sustainable Resource Science, Hirosawa, Wako, Saitama, Japan

Naoshi Dohmae

Roles Formal analysis, Investigation

Affiliation Biomolecular Characterization Unit, RIKEN Center for Sustainable Resource Science, Hirosawa, Wako, Saitama, Japan

Kazuya Takizawa

Roles Formal analysis, Investigation

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Yuka Nakazawa

Roles Investigation

Affiliations Department of Genetics, Research Institute of Environmental Medicine, Nagoya University, Furo-cho, Chikusa-ku, Nagoya, Japan, Department of Genome Repair, Atomic Bomb Disease Institute, Nagasaki University, Sakamoto, Nagasaki, Japan

Matthew D. Genet

Roles Investigation

Affiliation Department of Environmental and Radiological Health Sciences, Colorado State University, Fort Collins, CO, United States of America

Mika Saotome

Roles Investigation

Affiliation Program in Chemistry and Life Science, Department of Interdisciplinary Science and Engineering, School of Science and Engineering, Meisei University, Hodokubo, Hino-shi, Tokyo, Japan

Michio Hama

Roles Investigation

Affiliation Department of Basic Medical Sciences for Radiation Damage, NIRS, National Institutes for Quantum and Radiation Sciences and Technology (QST), Anagawa, Inage-ku, Chiba, Japan

Teruaki Konishi

Roles Investigation, Methodology

Affiliation Department of Basic Medical Sciences for Radiation Damage, NIRS, National Institutes for Quantum and Radiation Sciences and Technology (QST), Anagawa, Inage-ku, Chiba, Japan

Nakako Izumi Nakajima

Roles Investigation

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Charged Particle Therapy, NIRS, Anagawa, Inage-ku, Chiba, Japan

Masaharu Hazawa

Roles Investigation

Affiliations Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan, Cell-Bionomics Research Unit, Innovative Integrated Bio-Research Core, Institute for Frontier Science Initiative, Kanazawa University, Kakuma-machi, Kanazawa, Japan

Masanori Tomita

Roles Methodology

Affiliation Radiation Safety Research Center, Nuclear Technology Research Laboratory, Central Research Institute of Electric Power Industry, Iwado Kita, Komae-shi, Tokyo, Japan

Manabu Koike

Roles Formal analysis

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Protection, NIRS, 4-9-1 Anagawa, Inage-ku, Chiba, Japan

Katsuko Noshiro

Roles Investigation

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Kenichi Tomiyama

Roles Investigation

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Chizuka Obara

Roles Investigation

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Takaya Gotoh

Roles Investigation

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Ayako Ui

Roles Methodology

Affiliation Genome regulation and Molecular pharmacogenomics, School of Bioscience and Biotechnology, Tokyo University of Technology, Katakuramachi, Hachioji City, Tokyo, Japan

Akira Fujimori

Roles Investigation, Resources

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliations Research Center for Charged Particle Therapy, NIRS, Anagawa, Inage-ku, Chiba, Japan, International Open Laboratory (IOL), NIRS, Anagawa, Inage-ku, Chiba, Japan

Fumiaki Nakayama

Roles Project administration

Affiliation Department of Basic Medical Sciences for Radiation Damage, NIRS, National Institutes for Quantum and Radiation Sciences and Technology (QST), Anagawa, Inage-ku, Chiba, Japan

Fumio Hanaoka

Roles Supervision, Writing – review & editing

Affiliation Faculty of Science, Gakushuin University, Mejiro, Toshima-ku, Tokyo, Japan

Kaoru Sugasawa

Roles Methodology, Resources, Writing – review & editing

Affiliation Biosignal Research Center, and Graduate School of Science, Kobe University, Rokkodai-cho, Nada-ku, Kobe, Japan

Ryuichi Okayasu

Roles Funding acquisition, Supervision, Writing – review & editing

Affiliation International Open Laboratory (IOL), NIRS, Anagawa, Inage-ku, Chiba, Japan

Penny A. Jeggo

Roles Supervision, Writing – review & editing

Affiliations International Open Laboratory (IOL), NIRS, Anagawa, Inage-ku, Chiba, Japan, Genome Damage and Stability Centre, University of Sussex, Brighton, United Kingdom

Katsushi Tajima

Roles Supervision, Writing – original draft, Writing – review & editing

yasuda.takeshi@qst.go.jp (TY); tajima.katsushi@qst.go.jp (KT)

Current address: NIRS, QST, Anagawa, Inage-ku, Chiba, Japan

Affiliation Research Center for Radiation Emergency Medicine, National Institute of Radiological Sciences (NIRS), Anagawa, Inage-ku, Chiba, Japan

Competing Interests

The authors have declared that no competing interests exist.