About the Authors

Xingjie Hao

Roles Data curation, Methodology, Software, Visualization, Writing – original draft

Affiliations Key Laboratory of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education, Huazhong Agricultural University, Wuhan, Hubei, China, Department of Biostatistics, University of Michigan, Ann Arbor, MI, United States of America, Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, United States of America

Ping Zeng

Roles Methodology

Affiliations Department of Biostatistics, University of Michigan, Ann Arbor, MI, United States of America, Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, United States of America

Shujun Zhang

Roles Supervision

Affiliation Key Laboratory of Agricultural Animal Genetics, Breeding and Reproduction of Ministry of Education, Huazhong Agricultural University, Wuhan, Hubei, China

Xiang Zhou

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – review & editing

xzhousph@umich.edu

Affiliations Department of Biostatistics, University of Michigan, Ann Arbor, MI, United States of America, Center for Statistical Genetics, University of Michigan, Ann Arbor, MI, United States of America

Competing Interests

The authors have declared that no competing interests exist.