About the Authors

Baolu Shi

Contributed equally to this work with: Baolu Shi, Jiangyang Xue

Roles Conceptualization, Formal analysis, Investigation, Methodology

Affiliations State Key Laboratory of Reproductive Medicine, Nanjing Medical University, Nanjing, China, Department of Biomedical Sciences, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Jiangyang Xue

Contributed equally to this work with: Baolu Shi, Jiangyang Xue

Roles Conceptualization, Investigation, Methodology

Affiliations State Key Laboratory of Reproductive Medicine, Nanjing Medical University, Nanjing, China, Department of Biomedical Sciences, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America, Center for Reproduction and Genetics, The Affiliated Suzhou Hospital of Nanjing Medical University, Suzhou, Jiangsu, China

Jian Zhou

Roles Conceptualization, Investigation, Methodology

Affiliation Xinhua Hospital, Affiliated to Shanghai Jiao Tong University School of Medicine, Shanghai, China

Seth D. Kasowitz

Roles Data curation, Formal analysis

Affiliation Department of Biomedical Sciences, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Yuanwei Zhang

Roles Formal analysis

Affiliation USTC-SJH Joint Center for Human Reproduction and Genetics, School of Life Sciences, University of Science and Technology of China, Hefei,Anhui, China

Guanxiang Liang

Roles Formal analysis

Affiliation Department of Microbiology, University of Pennsylvania Perelman School of Medicine, Philadelphia, Pennsylvania, United States of America

Yongjuan Guan

Roles Investigation, Resources

Affiliation Department of Biomedical Sciences, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Qinghua Shi

Roles Formal analysis

Affiliation USTC-SJH Joint Center for Human Reproduction and Genetics, School of Life Sciences, University of Science and Technology of China, Hefei,Anhui, China

Mingxi Liu

Roles Supervision, Writing – review & editing

Affiliation State Key Laboratory of Reproductive Medicine, Nanjing Medical University, Nanjing, China

Jiahao Sha

Roles Funding acquisition, Investigation, Supervision, Writing – review & editing

Affiliation State Key Laboratory of Reproductive Medicine, Nanjing Medical University, Nanjing, China

Xiaoyan Huang

Roles Conceptualization, Funding acquisition, Investigation, Supervision, Writing – review & editing

bbhxy@njmu.edu.cn (XH); pwang@vet.upenn.edu (PJW)

Affiliation State Key Laboratory of Reproductive Medicine, Nanjing Medical University, Nanjing, China

P. Jeremy Wang

Roles Conceptualization, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

bbhxy@njmu.edu.cn (XH); pwang@vet.upenn.edu (PJW)

Affiliations State Key Laboratory of Reproductive Medicine, Nanjing Medical University, Nanjing, China, Department of Biomedical Sciences, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.