About the Authors

Sean C. Shadle

Affiliations Molecular and Cellular Biology Program, University of Washington, Seattle, Washington, United States of America, Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Jun Wen Zhong

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Amy E. Campbell

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Melissa L. Conerly

Affiliations Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Division of Basic Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Sujatha Jagannathan

Affiliations Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Division of Basic Sciences, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America, Computational Biology Program, Public Health Sciences Division, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Chao-Jen Wong

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Timothy D. Morello

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Silvère M. van der Maarel

Affiliation Department of Human Genetics, Leiden University Medical Center, Leiden, Netherlands

Stephen J. Tapscott

stapscot@fredhutch.org

Affiliation Division of Human Biology, Fred Hutchinson Cancer Research Center, Seattle, Washington, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: SCS SMvdM SJT. Data curation: CJW. Formal analysis: SCS CJW. Funding acquisition: SCS AEC SJ SJT. Investigation: SCS JWZ AEC MLC SJ TDM. Methodology: SCS JWZ SJT. Project administration: SCS SJT. Supervision: SCS SJT. Validation: SCS JWZ SJ AEC. Visualization: SCS. Writing – original draft: SCS SJT. Writing – review & editing: SCS JWZ AEC MLC SJ CJW SMvdM SJT.