About the Authors

Guangchao Liu

Contributed equally to this work with: Guangchao Liu, Shan Gao

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Shan Gao

Contributed equally to this work with: Guangchao Liu, Shan Gao

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Huiyu Tian

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Wenwen Wu

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Hélène S. Robert

Affiliation Mendel Centre for Genomics and Proteomics of Plants Systems, CEITEC MU—Central European Institute of Technology, Masaryk University, Brno, Czech Republic

Zhaojun Ding

dingzhaojun@sdu.edu.cn

Affiliation The Key Laboratory of Plant Cell Engineering and Germplasm Innovation, Ministry of Education, College of Life Science, Shandong University, Jinan, People’s Republic of China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceptualization: ZD. Formal analysis: GL SG HT. Funding acquisition: ZD HT. Investigation: GL SG WW HSR. Supervision: ZD HT. Writing – original draft: GL SG HSR HT ZD.