About the Authors

Kristen Fortney

Affiliations Department of Developmental Biology, Stanford University, Stanford, California, United States of America, Department of Genetics, Stanford University, Stanford, California, United States of America

Edgar Dobriban

Affiliation Department of Statistics, Stanford University, Stanford, California, United States of America

Paolo Garagnani

Affiliations Department of Experimental, Diagnostic and Specialty Medicine Experimental Pathology, University of Bologna, Bologna, Italy, Center for Applied Biomedical Research, St. Orsola-Malpighi University Hospital, Bologna, Italy

Chiara Pirazzini

Affiliations Department of Experimental, Diagnostic and Specialty Medicine Experimental Pathology, University of Bologna, Bologna, Italy, Interdepartmental Centre "L. Galvani" CIG, University of Bologna, Bologna, Italy

Daniela Monti

Affiliation Department of Clinical, Experimental and Biomedical Sciences, University of Florence, Florence, Italy

Daniela Mari

Affiliations Department of Medical Sciences, University of Milan, Milan, Italy, Geriatric Unit, IRCCS Ca' Grande Foundation, Maggiore Policlinico Hospital, Milan, Italy

Gil Atzmon

Affiliation Department of Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Nir Barzilai

Affiliation Department of Genetics, Albert Einstein College of Medicine, Bronx, New York, United States of America

Claudio Franceschi

Affiliations Department of Experimental, Diagnostic and Specialty Medicine Experimental Pathology, University of Bologna, Bologna, Italy, IRCCS, Institute of Neurological Sciences of Bologna, Bologna, Italy

Art B. Owen

Affiliation Department of Statistics, Stanford University, Stanford, California, United States of America

Stuart K. Kim

stuartkm@stanford.edu

Affiliations Department of Developmental Biology, Stanford University, Stanford, California, United States of America, Department of Genetics, Stanford University, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KF ED ABO SKK. Performed the experiments: KF ED PG CP. Analyzed the data: KF ED. Contributed reagents/materials/analysis tools: PG CP DMo DMa GA NB CF. Wrote the paper: KF SKK.