About the Authors

Cecil M. Benitez

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Kun Qu

Affiliation Program in Epithelial Biology, Stanford University School of Medicine, Stanford, California, United States of America

Takuya Sugiyama

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Philip T. Pauerstein

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Yinghua Liu

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Jennifer Tsai

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Xueying Gu

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Amar Ghodasara

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

H. Efsun Arda

Affiliation Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America

Jiajing Zhang

Affiliation Program in Epithelial Biology, Stanford University School of Medicine, Stanford, California, United States of America

Joseph D. Dekker

Affiliation Institute for Cellular and Molecular Biology and Department of Molecular Biosciences, University of Texas at Austin, Austin, Texas, United States of America

Haley O. Tucker

Affiliation Institute for Cellular and Molecular Biology and Department of Molecular Biosciences, University of Texas at Austin, Austin, Texas, United States of America

Howard Y. Chang

Affiliations Program in Epithelial Biology, Stanford University School of Medicine, Stanford, California, United States of America, Department of Dermatology, Stanford University School of Medicine,, Stanford, California, United States of America, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America

Seung K. Kim

seungkim@stanford.edu

Affiliations Department of Developmental Biology, Stanford University School of Medicine, Stanford, California, United States of America, Howard Hughes Medical Institute, Stanford University School of Medicine, Stanford, California, United States of America, Department of Medicine (Oncology Division), Stanford University School of Medicine, Stanford, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CMB TS SKK. Performed the experiments: CMB KQ TS PTP YL XG AG JZ JT. Analyzed the data: CMB SKK. Contributed reagents/materials/analysis tools: JDD HOT HYC HEA. Wrote the paper: CMB SKK.