About the Authors

Jessica Hopkins

Affiliation Department of Biochemistry and Molecular Biology, Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Grace Hwang

Contributed equally to this work with: Grace Hwang, Justin Jacob, Nicklas Sapp

Affiliation Department of Biochemistry and Molecular Biology, Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Justin Jacob

Contributed equally to this work with: Grace Hwang, Justin Jacob, Nicklas Sapp

Affiliation Department of Biochemistry and Molecular Biology, Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Nicklas Sapp

Contributed equally to this work with: Grace Hwang, Justin Jacob, Nicklas Sapp

Affiliation Department of Biochemistry and Molecular Biology, Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Rick Bedigian

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Kazuhiro Oka

Affiliation Department of Molecular and Cellular Biology, Baylor College of Medicine, Houston, Texas, United States of America, Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America

Paul Overbeek

Affiliation Department of Molecular and Cellular Biology, Baylor College of Medicine, Houston, Texas, United States of America, Department of Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas, United States of America

Steve Murray

Affiliation The Jackson Laboratory, Bar Harbor, Maine, United States of America

Philip W. Jordan

pjordan8@jhu.edu

Affiliation Department of Biochemistry and Molecular Biology, Johns Hopkins University Bloomberg School of Public Health, Baltimore, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JH GH JJ NS PWJ. Performed the experiments: JH GH JJ NS PWJ. Analyzed the data: JH GH JJ NS PWJ. Contributed reagents/materials/analysis tools: RB SM PO KO. Wrote the paper: JH GH PWJ.