About the Authors

Adam H. Freedman

Affiliation Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, California, United States of America

Ilan Gronau

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Rena M. Schweizer

Affiliation Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, California, United States of America

Diego Ortega-Del Vecchyo

Affiliation Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, California, United States of America

Eunjung Han

Affiliation Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, California, United States of America

Pedro M. Silva

Affiliation CIBIO-UP, University of Porto, Vairão, Portugal

Marco Galaverni

Affiliation ISPRA, Ozzano dell'Emilia, Italy

Zhenxin Fan

Affiliation Key Laboratory of Bioresources and Ecoenvironment, Sichuan University, Chengdu, China

Peter Marx

Affiliation Department of Measurement and Information Systems, Budapest University of Technology and Economics, Budapest, Hungary

Belen Lorente-Galdos

Affiliation Institut de Biologia Evolutiva (CSIC-Univ Pompeu Fabra), Barcelona, Spain

Holly Beale

Affiliation National Institutes of Health/NHGRI, Bethesda, Maryland, United States of America

Oscar Ramirez

Affiliation Institut de Biologia Evolutiva (CSIC-Univ Pompeu Fabra), Barcelona, Spain

Farhad Hormozdiari

Affiliation Department of Computer Science, University of California, Los Angeles, Los Angeles, California, United States of America

Can Alkan

Affiliation Bilkent University, Ankara, Turkey

Carles Vilà

Affiliation Estación Biológia de Doñana EBD-CSIC, Sevilla, Spain

Kevin Squire

Affiliation Department of Human Genetics, University of California, Los Angeles, Los Angeles, California, United States of America

Eli Geffen

Affiliation Department of Zoology, Tel Aviv University, Tel Aviv, Israel

Josip Kusak

Affiliation University of Zagreb, Zagreb, Croatia

Adam R. Boyko

Affiliation Department of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Heidi G. Parker

Affiliation National Institutes of Health/NHGRI, Bethesda, Maryland, United States of America

Clarence Lee

Affiliation Life Technologies, Foster City, California, United States of America

Vasisht Tadigotla

Affiliation Life Technologies, Foster City, California, United States of America

Adam Siepel

Affiliation Department of Biological Statistics and Computational Biology, Cornell University, Ithaca, New York, United States of America

Carlos D. Bustamante

Affiliation Stanford School of Medicine, Stanford, California, United States of America

Timothy T. Harkins

Affiliation Life Technologies, Foster City, California, United States of America

Stanley F. Nelson

Affiliation Department of Human Genetics, University of California, Los Angeles, Los Angeles, California, United States of America

Elaine A. Ostrander

Affiliation National Institutes of Health/NHGRI, Bethesda, Maryland, United States of America

Tomas Marques-Bonet

Affiliations Institut de Biologia Evolutiva (CSIC-Univ Pompeu Fabra), Barcelona, Spain, Institució Catalana de Recerca i Estudis Avançats (ICREA). 08010, Barcelona, Spain

Robert K. Wayne

rwayne@ucla.edu (RKW); jnovembre@uchicago.edu (JN)

Affiliation Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, California, United States of America

John Novembre

rwayne@ucla.edu (RKW); jnovembre@uchicago.edu (JN)

Current address: Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America.

Affiliation Department of Ecology and Evolutionary Biology, University of California, Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: AHF CDB TTH SFN EAO TMB RKW JN. Performed the experiments: RMS BLG HB OR CV KS ARB HGP CL VT. Analyzed the data: AHF RMS IG EH DODV PMS MG ZF PM BLG OR FH CA VT AS JN. Contributed reagents/materials/analysis tools: CV EG JK EAO RKW. Wrote the paper: AHF RMS IG EH DODV PMS MG ZF PM HB AS TMB RKW JN.