About the Authors

Maxim V. C. Greenberg

Contributed equally to this work with: Maxim V. C. Greenberg, Angelique Deleris, Christopher J. Hale

Current address: Unité de Génétique et Biologie du Développement, Institut Curie, Paris, France.

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Angelique Deleris

Contributed equally to this work with: Maxim V. C. Greenberg, Angelique Deleris, Christopher J. Hale

Current address: Section de Génomique Environnementale et Evolutive, Institut de Biologie de l'Ecole Normale Supérieure, Paris, France.

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Christopher J. Hale

Contributed equally to this work with: Maxim V. C. Greenberg, Angelique Deleris, Christopher J. Hale

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Ao Liu

Affiliation Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America

Suhua Feng

suhuaf@ucla.edu (SF); jacobsen@ucla.edu (SEJ)

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Eli & Edythe Broad Center of Regenerative Medicine & Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America, Howard Hughes Medical Institute, University of California Los Angeles, Los Angeles, California, United States of America

Steven E. Jacobsen

suhuaf@ucla.edu (SF); jacobsen@ucla.edu (SEJ)

Affiliations Department of Molecular, Cell and Developmental Biology, University of California Los Angeles, Los Angeles, California, United States of America, Eli & Edythe Broad Center of Regenerative Medicine & Stem Cell Research, University of California Los Angeles, Los Angeles, California, United States of America, Howard Hughes Medical Institute, University of California Los Angeles, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MVCG AD CJH SEJ. Performed the experiments: MVCG AD CJH AL SF. Analyzed the data: MVCG AD CJH SEJ. Wrote the paper: MVCG AD CJH SF SEJ.