About the Authors

Martin Ladouceur

Affiliations Department of Human Genetics, McGill University, Montreal, Canada, Lady Davis Institute for Medical Research, Jewish General Hospital, Montreal, Canada

Zari Dastani

Affiliations Lady Davis Institute for Medical Research, Jewish General Hospital, Montreal, Canada, Department of Epidemiology, Biostatistics and Occupational Health, McGill University, Montreal, Canada

Yurii S. Aulchenko

Affiliations Department of Epidemiology, Erasmus MC, Rotterdam, The Netherlands, Institute of Cytology and Genetics SD RAS, Novosibirsk, Russia

Celia M. T. Greenwood

Affiliations Lady Davis Institute for Medical Research, Jewish General Hospital, Montreal, Canada, Department of Epidemiology, Biostatistics and Occupational Health, McGill University, Montreal, Canada, Department of Oncology, McGill University, Montreal, Canada

J. Brent Richards

brent.richards@mcgill.ca

Affiliations Department of Human Genetics, McGill University, Montreal, Canada, Lady Davis Institute for Medical Research, Jewish General Hospital, Montreal, Canada, Department of Medicine, Jewish General Hospital, McGill University, Montreal, Canada, Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ML YSA CMTG JBR. Performed the experiments: ML CMTG. Analyzed the data: ML. Contributed reagents/materials/analysis tools: ML ZD YSA CMTG JBR. Wrote the paper: ML ZD YSA CMTG JBR. Designed the software used in analysis: ML CMTG. Developed the WOD algorithm: CMTG. Obtained permission for the data: JBR. Performed statistical analysis and simulations: ML.