About the Authors

Molly E. McCue

mccu0173@umn.edu

Affiliation College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Danika L. Bannasch

Affiliation Department of Population Health and Reproduction, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Jessica L. Petersen

Affiliation College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Jessica Gurr

Affiliation Faculty of Veterinary Science, University of Sydney, Sydney, Australia

Ernie Bailey

Affiliation Maxwell H. Gluck Equine Research Center, Department of Veterinary Science, University of Kentucky, Lexington, Kentucky, United States of America

Matthew M. Binns

Affiliation Equine Analysis Systems, Midway, Kentucky, United States of America

Ottmar Distl

Affiliation Institute of Animal Breeding and Genetics, University of Veterinary Medicine Hannover, Hannover, Germany

Gérard Guérin

Affiliation INRA, UMR 1313, Génétique Animale et Biologie Intégrative, Biologie Intégrative et Génétique Equine, Jouy-en-Josas, France

Telhisa Hasegawa

Affiliation Equine Research Institute, Japan Racing Association, Utsunomiya, Japan

Emmeline W. Hill

Affiliation Animal Genomics Laboratory, School of Agriculture and Food Science, University College Dublin, Dublin, Ireland

Tosso Leeb

Affiliation Institute of Genetics, Vetsuisse Faculty, University of Bern, Berne, Switzerland

Gabriella Lindgren

Affiliation Department of Animal Breeding and Genetics, Swedish University of Agricultural Sciences, Uppsala, Sweden

M. Cecilia T. Penedo

Affiliation Veterinary Genetics Laboratory, University of California Davis, Davis, California, United States of America

Knut H. Røed

Affiliation Department of Basic Sciences and Aquatic Medicine, Norwegian School of Veterinary Science, Oslo, Norway

Oliver A. Ryder

Affiliation San Diego Zoo's Institute for Conservation Research, Escondido, California, United States of America

June E. Swinburne

Affiliation Animal Health Trust, Suffolk, United Kingdom

Teruaki Tozaki

Affiliation Department of Molecular Genetics, Laboratory of Racing Chemistry, Utsunomiya, Japan

Stephanie J. Valberg

Affiliation College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Mark Vaudin

Affiliation Animal Health Trust, Suffolk, United Kingdom

Kerstin Lindblad-Toh

Affiliations Science for Life Laboratory, Department of Medical Biochemistry and Microbiology, Uppsala University, Uppsala, Sweden, Broad Institute, Cambridge, Massachusetts, United States of America

Claire M. Wade

Contributed equally to this work with: Claire M. Wade, James R. Mickelson

Affiliation Faculty of Veterinary Science, University of Sydney, Sydney, Australia

James R. Mickelson

Contributed equally to this work with: Claire M. Wade, James R. Mickelson

Affiliation College of Veterinary Medicine, University of Minnesota, St. Paul, Minnesota, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MEM DLB MMB KL-T CMW JRM OAR. Analyzed the data: MEM JLP JG DLB CMW. Contributed reagents/materials/analysis tools: EB MMB OD GG TH EWH TL GL MCTP KHR OAR JES TT SJV MV. Wrote the paper: MEM JLP JRM.