About the Authors

Xavier Didelot

Contributed equally to this work with: Xavier Didelot, Rory Bowden

Affiliation Department of Statistics, Oxford University, Oxford, United Kingdom

Rory Bowden

Contributed equally to this work with: Xavier Didelot, Rory Bowden

Affiliations Department of Statistics, Oxford University, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, Oxford University, Oxford, United Kingdom

Teresa Street

Affiliation Department of Statistics, Oxford University, Oxford, United Kingdom

Tanya Golubchik

Affiliation Department of Statistics, Oxford University, Oxford, United Kingdom

Chris Spencer

Affiliation Wellcome Trust Centre for Human Genetics, Oxford University, Oxford, United Kingdom

Gil McVean

Affiliations Department of Statistics, Oxford University, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, Oxford University, Oxford, United Kingdom

Vartul Sangal

Affiliation Strathclyde Institute of Pharmacy and Biomedical Sciences, University of Strathclyde, Glasgow, United Kingdom

Muna F. Anjum

Affiliation Veterinary Laboratories Agency, Addlestone, United Kingdom

Mark Achtman

Affiliation Environmental Research Institute and Department of Microbiology, University College Cork, Cork, Ireland

Daniel Falush

Affiliations Environmental Research Institute and Department of Microbiology, University College Cork, Cork, Ireland, Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany

Peter Donnelly

donnelly@stats.ox.ac.uk

Affiliations Department of Statistics, Oxford University, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, Oxford University, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XD RB GM DF PD. Performed the experiments: RB TS. Analyzed the data: XD TG CS. Contributed reagents/materials/analysis tools: VS MFA MA. Wrote the paper: XD RB DF PD.