About the Authors

Robert S. Illingworth

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Ulrike Gruenewald-Schneider

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Shaun Webb

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Alastair R. W. Kerr

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Keith D. James

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Daniel J. Turner

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Colin Smith

Affiliation Neuropathology Unit, Division of Pathology, University of Edinburgh, Edinburgh, United Kingdom

David J. Harrison

Affiliation Division of Pathology, Institute of Genetics and Molecular Medicine, University of Edinburgh, Western General Hospital, Edinburgh, United Kingdom

Robert Andrews

Affiliation Wellcome Trust Sanger Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Adrian P. Bird

a.bird@ed.ac.uk

Affiliation Wellcome Trust Centre for Cell Biology, University of Edinburgh, Edinburgh, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RSI APB. Performed the experiments: RSI UGS DJT. Analyzed the data: RSI UGS SW ARWK KDJ DJT RA. Contributed reagents/materials/analysis tools: CS DJH. Wrote the paper: RSI APB.