About the Authors

Fan Liu

Affiliation Department of Forensic Molecular Biology, Erasmus University Medical Center, Rotterdam, The Netherlands

Andreas Wollstein

Affiliations Department of Forensic Molecular Biology, Erasmus University Medical Center, Rotterdam, The Netherlands, Cologne Center for Genomics (CCG), University of Cologne, Cologne, Germany

Pirro G. Hysi

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Georgina A. Ankra-Badu

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Timothy D. Spector

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Daniel Park

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Gu Zhu

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Mats Larsson

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

David L. Duffy

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Grant W. Montgomery

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

David A. Mackey

Affiliation Centre for Ophthalmology and Visual Science, Lions Eye Institute, University of Western Australia, Perth, Australia

Susan Walsh

Affiliation Department of Forensic Molecular Biology, Erasmus University Medical Center, Rotterdam, The Netherlands

Oscar Lao

Affiliation Department of Forensic Molecular Biology, Erasmus University Medical Center, Rotterdam, The Netherlands

Albert Hofman

Affiliation Department of Epidemiology, Erasmus University Medical Center, Rotterdam, The Netherlands

Fernando Rivadeneira

Affiliations Department of Epidemiology, Erasmus University Medical Center, Rotterdam, The Netherlands, Department of Internal Medicine, Erasmus University Medical Center, Rotterdam, The Netherlands

Johannes R. Vingerling

Affiliations Department of Epidemiology, Erasmus University Medical Center, Rotterdam, The Netherlands, Department of Ophthalmology, Erasmus University Medical Center, Rotterdam, The Netherlands

André G. Uitterlinden

Affiliations Department of Epidemiology, Erasmus University Medical Center, Rotterdam, The Netherlands, Department of Internal Medicine, Erasmus University Medical Center, Rotterdam, The Netherlands

Nicholas G. Martin

Affiliation Queensland Institute of Medical Research, Brisbane, Australia

Christopher J. Hammond

Affiliation Department of Twin Research and Genetic Epidemiology, King's College London, London, United Kingdom

Manfred Kayser

m.kayser@erasmusmc.nl

Affiliation Department of Forensic Molecular Biology, Erasmus University Medical Center, Rotterdam, The Netherlands

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FL AW JRV AGU NGM CJH MK. Performed the experiments: FL AW PH GAAB DP GZ ML DLD GWM SW OL FR MK. Analyzed the data: FL AW PH GAAB DP GZ ML DLD GWM SW OL FR MK. Contributed reagents/materials/analysis tools: TDS DAM AH FR JRV AGU NGM CJH MK. Wrote the paper: FL AW PH GAAB TDS SW OL AH FR JRV AGU NGM CJH MK.