About the Authors

Bryan N. Howie

Current address: Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Peter Donnelly

Affiliations Department of Statistics, University of Oxford, Oxford, United Kingdom, Wellcome Trust Centre for Human Genetics, Roosevelt Drive, Oxford, United Kingdom

Jonathan Marchini

marchini@stats.ox.ac.uk

Affiliation Department of Statistics, University of Oxford, Oxford, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BNH JM. Performed the experiments: BNH. Analyzed the data: BNH. Wrote the paper: BNH PD JM.