About the Authors

Filippo Del Bene

Contributed equally to this work with: Filippo Del Bene, Laurence Ettwiller

Affiliations Developmental Biology Programme, European Molecular Biology Laboratory, Heidelberg, Heidelberg, Germany , Department of Physiology, University of California San Francisco, San Francisco, California, United States of America , Programs in Neuroscience, Genetics, and Developmental Biology, University of California San Francisco, San Francisco, California, United States of America , Center for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Laurence Ettwiller

Contributed equally to this work with: Filippo Del Bene, Laurence Ettwiller

Affiliations Developmental Biology Programme, European Molecular Biology Laboratory, Heidelberg, Heidelberg, Germany , European Bioinformatics Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom , Neurobiologie et diversité cellulaire, Ecole Supérieure de Physique et de Chimie Industrielles, CNRS, UMR7637, Paris, France

Dorota Skowronska-Krawczyk

Affiliation University of Lausanne, Eye Hospital Jules Gonin, Lausanne, Switzerland

Herwig Baier

Affiliations Department of Physiology, University of California San Francisco, San Francisco, California, United States of America , Programs in Neuroscience, Genetics, and Developmental Biology, University of California San Francisco, San Francisco, California, United States of America , Center for Human Genetics, University of California San Francisco, San Francisco, California, United States of America

Jean-Marc Matter

Affiliation University of Lausanne, Eye Hospital Jules Gonin, Lausanne, Switzerland

Ewan Birney

To whom correspondence should be addressed. E-mail: Jochen.Wittbrodt@embl.de (JW); (birney@ebi.ac.uk (EB)

Affiliation European Bioinformatics Institute, Wellcome Trust Genome Campus, Hinxton, Cambridge, United Kingdom

Joachim Wittbrodt

To whom correspondence should be addressed. E-mail: Jochen.Wittbrodt@embl.de (JW); (birney@ebi.ac.uk (EB)

Affiliation Developmental Biology Programme, European Molecular Biology Laboratory, Heidelberg, Heidelberg, Germany

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

FDB, LE, and JW conceived and designed the experiments and wrote the paper. FDB, LE, and DSK performed the experiments. FDB, LE, JMM, EB, and JW analyzed the data. FDB, LE, DSK, HB, and JMM contributed reagents/materials/analysis tools.