About the Authors

Celine Becquet

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

Nick Patterson

Affiliation Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America

Anne C Stone

Affiliation School of Human Evolution and Social Change, Arizona State University, Tempe, Arizona, United States of America

Molly Przeworski

To whom correspondence should be addressed. E-mail: mfp@uchicago.edu (MP); reich@genetics.med.harvard.edu (DR)

Affiliation Department of Human Genetics, University of Chicago, Chicago, Illinois, United States of America

David Reich

To whom correspondence should be addressed. E-mail: mfp@uchicago.edu (MP); reich@genetics.med.harvard.edu (DR)

Affiliations Broad Institute of Harvard and MIT, Cambridge, Massachusetts, United States of America , Department of Genetics, Harvard Medical School, Boston, Massachusetts, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

MP and DR conceived and designed the experiments. CB, MP, and DR performed the experiments and wrote the paper. CB, NP, MP, and DR analyzed the data. CB, NP, ACS, MP, and DR contributed reagents/materials/analysis tools. MP and DR cosupervised the work.