A positive-sense single-stranded RNA virus acquired a negative-sense open reading frame through recombination

• Xuan Dong ,

  Contributed equally to this work with: Xuan Dong, Fan Zhang

  Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Software, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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• Fan Zhang ,

  Contributed equally to this work with: Xuan Dong, Fan Zhang

  Roles Data curation, Formal analysis, Investigation, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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• Yiting Wang,

  Roles Conceptualization, Data curation, Formal analysis, Investigation, Software, Writing – review & editing

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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• Cixiu Li,

  Roles Data curation, Formal analysis, Software, Writing – review & editing

  Affiliation Key Laboratory of Emerging Infectious Diseases in Universities of Shandong, Shandong First Medical University & Shandong Academy of Medical Sciences, Ji’nan, China

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• Xuan Li,

  Roles Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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• Chengyan Zhou,

  Roles Data curation, Formal analysis, Software, Visualization, Writing – original draft, Writing – review & editing

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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• Guohao Wang,

  Roles Investigation, Validation, Writing – review & editing

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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• Zhongshuai Tian,

  Roles Formal analysis, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

  Affiliation Shanghai Institute of Virology, Shanghai Jiao Tong University School of Medicine, Shanghai, China

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• Jing Gao,

  Roles Formal analysis, Investigation, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

  Affiliation Department of Analytical Chemistry, State Key Laboratory of Drug Research, Shanghai Institute of Materia Medica, Chinese Academy of Sciences, Shanghai, China

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• Hu Zhou,

  Roles Methodology, Supervision, Writing – review & editing

  Affiliation Department of Analytical Chemistry, State Key Laboratory of Drug Research, Shanghai Institute of Materia Medica, Chinese Academy of Sciences, Shanghai, China

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• Liying Sui,

  Roles Resources, Supervision, Writing – review & editing

  Affiliation Asian Regional Artemia Reference Center, Tianjin University of Science & Technology, Tianjin, China

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• Hans Nauwynck,

  Roles Resources, Supervision, Writing – review & editing

  Affiliation Laboratory of Virology, Faculty of Veterinary Medicine, Ghent University, Ghent, Belgium

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• Patrick Sorgeloos,

  Roles Resources, Supervision, Writing – review & editing

  Affiliation Laboratory of Aquaculture & Artemia Reference Center, Faculty of Bioscience Engineering, Ghent University, Ghent, Belgium

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• Edward C. Holmes,

  Roles Methodology, Supervision, Writing – review & editing

  Affiliation School of Medical Sciences, The University of Sydney, Sydney, Australia

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• Jie Huang ,

  Roles Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – review & editing

  * E-mail: huangjie@ysfri.ac.cn (JH); shiwf@ioz.ac.cn (WS)

  Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

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•  [ ... ],
• Weifeng Shi

  Roles Conceptualization, Funding acquisition, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

  * E-mail: huangjie@ysfri.ac.cn (JH); shiwf@ioz.ac.cn (WS)

  Affiliations Shanghai Institute of Virology, Shanghai Jiao Tong University School of Medicine, Shanghai, China, Ruijin Hospital, Shanghai Jiao Tong University School of Medicine, Shanghai, China

  https://orcid.org/0000-0002-8717-2942

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