About the Authors

Xuan Dong

Contributed equally to this work with: Xuan Dong, Fan Zhang

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Investigation, Methodology, Project administration, Software, Supervision, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Fan Zhang

Contributed equally to this work with: Xuan Dong, Fan Zhang

Roles Data curation, Formal analysis, Investigation, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Yiting Wang

Roles Conceptualization, Data curation, Formal analysis, Investigation, Software, Writing – review & editing

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Cixiu Li

Roles Data curation, Formal analysis, Software, Writing – review & editing

Affiliation Key Laboratory of Emerging Infectious Diseases in Universities of Shandong, Shandong First Medical University & Shandong Academy of Medical Sciences, Ji’nan, China

Xuan Li

Roles Formal analysis, Investigation, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Chengyan Zhou

Roles Data curation, Formal analysis, Software, Visualization, Writing – original draft, Writing – review & editing

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Guohao Wang

Roles Investigation, Validation, Writing – review & editing

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Zhongshuai Tian

Roles Formal analysis, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Shanghai Institute of Virology, Shanghai Jiao Tong University School of Medicine, Shanghai, China

Jing Gao

Roles Formal analysis, Investigation, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Analytical Chemistry, State Key Laboratory of Drug Research, Shanghai Institute of Materia Medica, Chinese Academy of Sciences, Shanghai, China

Hu Zhou

Roles Methodology, Supervision, Writing – review & editing

Affiliation Department of Analytical Chemistry, State Key Laboratory of Drug Research, Shanghai Institute of Materia Medica, Chinese Academy of Sciences, Shanghai, China

Liying Sui

Roles Resources, Supervision, Writing – review & editing

Affiliation Asian Regional Artemia Reference Center, Tianjin University of Science & Technology, Tianjin, China

Hans Nauwynck

Roles Resources, Supervision, Writing – review & editing

Affiliation Laboratory of Virology, Faculty of Veterinary Medicine, Ghent University, Ghent, Belgium

Patrick Sorgeloos

Roles Resources, Supervision, Writing – review & editing

Affiliation Laboratory of Aquaculture & Artemia Reference Center, Faculty of Bioscience Engineering, Ghent University, Ghent, Belgium

Edward C. Holmes

Roles Methodology, Supervision, Writing – review & editing

Affiliation School of Medical Sciences, The University of Sydney, Sydney, Australia

Jie Huang

Roles Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – review & editing

huangjie@ysfri.ac.cn (JH); shiwf@ioz.ac.cn (WS)

Affiliation State Key Laboratory of Mariculture Biobreeding and Sustainable Goods, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences; Laboratory for Marine Fisheries Science and Food Production Processes, Qingdao Marine Science and Technology Center; Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao, China

Weifeng Shi

Roles Conceptualization, Funding acquisition, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

huangjie@ysfri.ac.cn (JH); shiwf@ioz.ac.cn (WS)

Affiliations Shanghai Institute of Virology, Shanghai Jiao Tong University School of Medicine, Shanghai, China, Ruijin Hospital, Shanghai Jiao Tong University School of Medicine, Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.