About the Authors

Senzhao Zhang

Contributed equally to this work with: Senzhao Zhang, Xuedan Tang, Zhen Wu

Roles Conceptualization, Investigation, Methodology, Project administration, Visualization, Writing – original draft

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China

Xuedan Tang

Contributed equally to this work with: Senzhao Zhang, Xuedan Tang, Zhen Wu

Roles Investigation

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China

Zhen Wu

Contributed equally to this work with: Senzhao Zhang, Xuedan Tang, Zhen Wu

Roles Funding acquisition, Methodology, Supervision, Writing – review & editing

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Key Laboratory of Agricultural Bioinformatics, Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Hantai Tan

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China

Yan Zhang

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China

Wangyang Tan

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China

Yu He

Roles Formal analysis, Funding acquisition

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Key Laboratory of Agricultural Bioinformatics, Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Tao Wang

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Mingshu Wang

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Renyong Jia

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Dekang Zhu

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Mafeng Liu

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Xinxin Zhao

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Qiao Yang

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Ying Wu

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Shaqiu Zhang

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Juan Huang

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Xumin Ou

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Di Sun

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Bin Tian

Roles Formal analysis

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Anchun Cheng

Roles Funding acquisition, Resources

chenganchun@vip.163.com (AC), shunchen@sicau.edu.cn (SC)

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China, Institute of Veterinary Immunology and Green Drugs, Veterinary Department in College of Animal Science, State Key Laboratory of Green Pesticide, Guizhou University, Guiyang, China

Shun Chen

Roles Conceptualization, Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

chenganchun@vip.163.com (AC), shunchen@sicau.edu.cn (SC)

Affiliations Institute of Veterinary Medicine and Immunology, Sichuan Agricultural University, Chengdu, Sichuan, China, Research Center of Avian Disease, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, Sichuan, China, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Sichuan Agricultural University, Chengdu, Sichuan, China, Key Laboratory of Agricultural Bioinformatics, Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China, Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education, Sichuan Agricultural University, Chengdu, Sichuan, China

Competing Interests

The authors have declared that no competing interests exist.