About the Authors

Hui Yan

Roles Data curation, Formal analysis, Investigation, Methodology, Resources

‡ These authors contributed equally to this work.

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Lei Bei

Roles Investigation, Resources, Validation

‡ These authors contributed equally to this work.

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Mingrui Zhao

Roles Investigation, Validation

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Jiajun Wang

Roles Visualization

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Pengxiang Jin

Roles Resources

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Chunmei Xu

Roles Investigation

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Lin Ju

Roles Visualization

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Yu Meng

Roles Project administration

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Shijin Jiang

Roles Funding acquisition, Project administration, Supervision

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Ruihua Zhang

Roles Conceptualization, Data curation, Formal analysis, Funding acquisition, Methodology, Project administration, Supervision, Writing – original draft, Writing – review & editing

rhzhang@sdau.edu.cn

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Tai’an, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Tai’an, China

Competing Interests

The authors have declared that no competing interests exist.