About the Authors

Xiaolan Liu

Contributed equally to this work with: Xiaolan Liu, Ruihao Zhang, Anchun Cheng

Roles Data curation, Writing – original draft, Writing – review & editing

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Ruihao Zhang

Contributed equally to this work with: Xiaolan Liu, Ruihao Zhang, Anchun Cheng

Roles Investigation, Methodology

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Mingshu Wang

Roles Data curation, Writing – review & editing

2878665974@qq.com

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Anchun Cheng

Contributed equally to this work with: Xiaolan Liu, Ruihao Zhang, Anchun Cheng

Roles Funding acquisition, Project administration

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Wei Zhang

Roles Resources

Affiliation Sinopharm Yangzhou VAC Biological Engineering Co., Ltd., Yangzhou, China

Qiao Yang

Roles Project administration, Software

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Xumin Ou

Roles Data curation, Formal analysis

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Di Sun

Roles Conceptualization, Data curation

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Yu He

Roles Software, Supervision

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Bin Tian

Roles Supervision, Validation

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Zhen Wu

Roles Data curation, Visualization

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Shaqiu Zhang

Roles Data curation, Validation

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Juan Huang

Roles Supervision, Validation

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Ying Wu

Roles Data curation, Supervision

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Yanling Yu

Roles Data curation, Formal analysis

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Ling Zhang

Roles Resources, Supervision

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

XinXin Zhao

Roles Data curation, Investigation

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Dekang Zhu

Roles Resources, Supervision

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Shun Chen

Roles Data curation, Methodology

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Renyong Jia

Roles Investigation, Resources

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Mafeng Liu

Roles Resources, Supervision

Affiliation Engineering Research Center of Southwest Animal Disease Prevention and Control Technology for Ministry of Education of the People’s Republic of China, International Joint Research Center for Animal Disease Prevention and Control of Sichuan Province, Agricultural Animal Diseases and Veterinary Public Health Key Laboratory of Sichuan Province, Research Center of Avian Disease and Institute of Veterinary Medicine and Immunology, College of Veterinary Medicine, Sichuan Agricultural University, Chengdu, China

Competing Interests

The authors have declared that no competing interests exist.