About the Authors

Hejiao Zhao

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Taian, Shandong, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Taian, Shandong, China

Yalu Zhang

Roles Investigation, Validation

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Taian, Shandong, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Taian, Shandong, China

Qingbing Han

Roles Investigation, Methodology, Validation

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Taian, Shandong, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Taian, Shandong, China

Huanan Li

Roles Investigation, Methodology

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Taian, Shandong, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Taian, Shandong, China

Wenjun Liu

Roles Resources, Writing – review & editing

Affiliations CAS Key Laboratory of Pathogenic Microbiology and Immunology, Institute of Microbiology, Chinese Academy of Sciences, Beijing, China, Savaid Medical School, University of Chinese Academy of Sciences, Beijing, China, Shenzhen Medical Academy of Research and Translation (SMART), Shenzhen, Guangdong, China

Lei Sun

Roles Funding acquisition, Project administration, Resources, Supervision, Writing – review & editing

shangyl@sdau.edu.cn (YS); sunlei362@im.ac.cn (LS)

Affiliations CAS Key Laboratory of Pathogenic Microbiology and Immunology, Institute of Microbiology, Chinese Academy of Sciences, Beijing, China, Savaid Medical School, University of Chinese Academy of Sciences, Beijing, China, Shandong Public Health Clinical Center, Jinan, Shandong, China

Yingli Shang

Roles Conceptualization, Data curation, Funding acquisition, Project administration, Supervision, Visualization, Writing – original draft, Writing – review & editing

shangyl@sdau.edu.cn (YS); sunlei362@im.ac.cn (LS)

Affiliations Department of Preventive Veterinary Medicine, College of Veterinary Medicine, Shandong Agricultural University, Taian, Shandong, China, Shandong Provincial Key Laboratory of Zoonoses, Shandong Agricultural University, Taian, Shandong, China, Institute of Immunology, Shandong Agricultural University, Taian, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.