About the Authors

Haixin Liu

Contributed equally to this work with: Haixin Liu, Haolun Tian

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Software, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliations College of Veterinary Medicine, Northwest A&F University, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Ministry of Education of the People’s Republic of China, Yangling, China, Key Laboratory of Ruminant Disease Prevention and Control (West), Ministry of Agriculture and Rural Affairs, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Universities of Shaanxi Province, Yangling, China

Haolun Tian

Contributed equally to this work with: Haixin Liu, Haolun Tian

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft

Affiliations College of Veterinary Medicine, Northwest A&F University, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Ministry of Education of the People’s Republic of China, Yangling, China, Key Laboratory of Ruminant Disease Prevention and Control (West), Ministry of Agriculture and Rural Affairs, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Universities of Shaanxi Province, Yangling, China

Pengcheng Hao

Roles Methodology

Affiliation College of Veterinary Medicine, Northwest A&F University, Yangling, China

Huimin Du

Roles Methodology

Affiliation College of Veterinary Medicine, Northwest A&F University, Yangling, China

Kun Wang

Roles Data curation

Affiliation College of Veterinary Medicine, Northwest A&F University, Yangling, China

Yudong Qiu

Roles Methodology

Affiliation College of Veterinary Medicine, Northwest A&F University, Yangling, China

Xiangrui Yin

Roles Investigation, Software

Affiliation College of Veterinary Medicine, Northwest A&F University, Yangling, China

Nana Wu

Roles Data curation, Investigation

Affiliation College of Veterinary Medicine, Northwest A&F University, Yangling, China

Qian Du

Roles Validation

Affiliations College of Veterinary Medicine, Northwest A&F University, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Ministry of Education of the People’s Republic of China, Yangling, China, Key Laboratory of Ruminant Disease Prevention and Control (West), Ministry of Agriculture and Rural Affairs, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Universities of Shaanxi Province, Yangling, China

Dewen Tong

Roles Conceptualization, Funding acquisition, Supervision, Writing – review & editing

dwtong@nwsuaf.edu.cn (DT); Yonghuang@nwsuaf.edu.cn (YH)

Affiliations College of Veterinary Medicine, Northwest A&F University, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Ministry of Education of the People’s Republic of China, Yangling, China, Key Laboratory of Ruminant Disease Prevention and Control (West), Ministry of Agriculture and Rural Affairs, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Universities of Shaanxi Province, Yangling, China

Yong Huang

Roles Conceptualization, Formal analysis, Funding acquisition, Project administration, Supervision, Writing – review & editing

dwtong@nwsuaf.edu.cn (DT); Yonghuang@nwsuaf.edu.cn (YH)

Affiliations College of Veterinary Medicine, Northwest A&F University, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Ministry of Education of the People’s Republic of China, Yangling, China, Key Laboratory of Ruminant Disease Prevention and Control (West), Ministry of Agriculture and Rural Affairs, Yangling, China, Engineering Research Center of Efficient New Vaccines for Animals, Universities of Shaanxi Province, Yangling, China

Competing Interests

The authors have declared that no competing interests exist.