About the Authors

Xiao Guo

Contributed equally to this work with: Xiao Guo, Youquan Xin, Zehui Tong

Roles Data curation, Formal analysis, Investigation, Methodology, Visualization, Writing – original draft

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Youquan Xin

Contributed equally to this work with: Xiao Guo, Youquan Xin, Zehui Tong

Roles Data curation, Funding acquisition, Methodology

Affiliation Key Laboratory for Plague Prevention and Control of Qinghai Province, Qinghai Institute for Endemic Disease Prevention and Control, Xining, China

Zehui Tong

Contributed equally to this work with: Xiao Guo, Youquan Xin, Zehui Tong

Roles Data curation, Formal analysis, Investigation, Methodology, Validation

Affiliation School of Basic Medical Sciences, Anhui Medical University Hefei, China

Shiyang Cao

Roles Methodology, Resources

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Yuan Zhang

Roles Funding acquisition, Methodology

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Gengshan Wu

Roles Data curation, Methodology

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Hongyan Chen

Roles Data curation, Formal analysis, Methodology

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Tong Wang

Roles Formal analysis, Methodology

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Yajun Song

Roles Formal analysis, Methodology, Resources

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Qingwen Zhang

Roles Resources

Affiliation Key Laboratory for Plague Prevention and Control of Qinghai Province, Qinghai Institute for Endemic Disease Prevention and Control, Xining, China

Ruifu Yang

Roles Methodology, Resources, Supervision

ruifuyang@gmail.com (RY); zmduams@163.com (ZD)

Affiliation State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology, Beijing, China

Zongmin Du

Roles Conceptualization, Formal analysis, Funding acquisition, Methodology, Project administration, Resources, Supervision, Writing – review & editing

ruifuyang@gmail.com (RY); zmduams@163.com (ZD)

Affiliations School of Basic Medical Sciences, Anhui Medical University Hefei, China, Key Laboratory for Plague Prevention and Control of Qinghai Province, Qinghai Institute for Endemic Disease Prevention and Control, Xining, China

Competing Interests

The authors have declared that no competing interests exist.