About the Authors

Jiandong Chen

Contributed equally to this work with: Jiandong Chen, Hyuntae Byun

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Hyuntae Byun

Contributed equally to this work with: Jiandong Chen, Hyuntae Byun

Roles Conceptualization, Data curation, Formal analysis, Investigation, Methodology, Writing – review & editing

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Qianxuan She

Roles Data curation, Formal analysis, Investigation, Methodology

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Zhi Liu

Roles Investigation, Methodology

Current address: Huazhong University of Science and Technology, Wuhan, China

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Karl-Gustav Ruggeberg

Roles Investigation, Methodology

Current address: CytoSorbents Medical Inc., Princeton, New Jersey, United States of America

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Qinqin Pu

Roles Investigation, Methodology

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

I-Ji Jung

Roles Investigation, Methodology

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Dehao Zhu

Roles Investigation, Methodology

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Mary R. Brockett

Roles Investigation, Writing – review & editing

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Ansel Hsiao

Roles Conceptualization, Data curation, Writing – review & editing

Affiliation Department of Microbiology & Plant Pathology, University of California Riverside, Riverside, California, United States of America

Jun Zhu

Roles Conceptualization, Data curation, Funding acquisition, Investigation, Methodology, Supervision, Writing – original draft, Writing – review & editing

junzhu@pennmedicine.upenn.edu

Affiliation Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.