About the Authors

Bonnie Douglas

Roles Conceptualization, Formal analysis, Investigation, Project administration, Resources, Validation, Visualization, Writing – original draft, Writing – review & editing

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Yun Wei

Roles Methodology, Writing – review & editing

Affiliation Department of Oncology and Inflammation, Amgen Research, South San Francisco, California, United States of America

Xinshe Li

Roles Investigation, Methodology, Resources

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Annabel Ferguson

Roles Data curation, Formal analysis

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Li-Yin Hung

Roles Investigation, Writing – review & editing

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Christopher Pastore

Roles Investigation, Resources, Writing – review & editing

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Jonathan R Kurtz

Roles Methodology

Affiliation Flagship Labs 72, Inc., Cambridge, Massachusetts, United States of America

James B. McLachlan

Roles Methodology, Writing – review & editing

Affiliation Department of Microbiology and Immunology, Tulane University School of Medicine, New Orleans, Louisiana, United States of America

Thomas J. Nolan

Roles Methodology, Resources, Writing – review & editing

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

James Lok

Roles Conceptualization, Funding acquisition, Investigation, Project administration, Resources, Supervision, Visualization, Writing – review & editing

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

De’Broski R. Herbert

Roles Conceptualization, Funding acquisition, Project administration, Resources, Supervision, Visualization, Writing – original draft, Writing – review & editing

debroski@vet.upenn.edu

Affiliation Department of Pathobiology, University of Pennsylvania School of Veterinary Medicine, Philadelphia, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.